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Cited article:

Protamines, not histones, are the predominant basic proteins in sperm nuclei of solitary ascidian tunicates

M. Chiva, F. Lafargue, E. Rosenberg and H. E. Kasinsky
Journal of Experimental Zoology 263 (3) 338 (1992)
https://doi.org/10.1002/jez.1402630314