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Cited article:

Hybrid proteins, heterosis and the origin of species—I. Unusual variation of polychaete hyalinoecia “nothing dehydrogenases” and of quail coturnix erythrocyte enzymes

Clyde Manwell and C.M. Ann Baker
Comparative Biochemistry and Physiology 28 (3) 1007 (1969)
https://doi.org/10.1016/0010-406X(69)90544-1